Professor, PhD, Invited PI of the Laboratory of Behavioural Neurodynamics, Head of MEG/EEG at the Center of Functionally Integrative Neuroscience, Aarhus University; Invited professor at the Institute of Cognitive Neuroscience Centre for Cognition & Decision Making at HSE University

Shtyrov Yury

Research interests

Yury Shtyrov has many years of international experience in studying human neurocognitive functions, in particular neurobiological foundations of the human speech and language function. His particular contribution to the science of language has been in uncovering early and automatic stages of language processing and in detailing the time course of spoken language comprehension in the brain. This work has to a large extent contributed to a dramatic change in our understanding of how the brain analyses speech, which has occurred in recent years. It shows how memory traces for linguistic elements in the brain can be probed using objective imaging tools, how they develop with learning, interact on different levels, as well as the interaction between the cognitive systems of language and attention. Most importantly, this work shows that these different processes occur rapidly and in parallel, something that was first met with disbelief but is now becoming generally accepted thanks to this and similar work.

In the Lab

Lab PI, the founder and inspirer of everything that happens in the Lab.


  1. Vukovic, Nikola; Shtyrov, Yury. Learning with the wave of the hand: kinematic and TMS evidence of primary motor cortex role in category-specific encoding of word meaning. NeuroImage, Vol. 202, 116179, 11.2019.
  2. Vasilyeva, Marina J.; Knyazeva, Veronika M.; Aleksandrov, Aleksander A.; Shtyrov, Yury. Neurophysiological Correlates of Fast Mapping of Novel Words in the Adult Brain. Frontiers in Human Neuroscience, Vol. 13, 304, 19.09.2019.
  3. Jensen, Mads; Hyder, Rasha; Shtyrov, Yury MVPA Analysis of Intertrial Phase Coherence of Neuromagnetic Responses to Words Reliably Classifies Multiple Levels of Language Processing in the Brain. eNeuro, Vol. 6, No. 4, 07.08.2019.
  4. Mkrtychian, Nadezhda; Blagovechtchenski, Evgeny; Kurmakaeva, Diana; Gnedykh, Dana; Kostromina, Svetlana; Shtyrov, Yury. Concrete vs. Abstract Semantics : From Mental Representations to Functional Brain Mapping. Frontiers in Human Neuroscience, Vol. 13, 267, 08.2019.
  5. Pokhoday, Mikhail Y.; Shtyrov, Yury Y.; Myachykov, Andriy. Effects of visual priming and event orientation on word order choice in Russian sentence production. Frontiers in Psychology, Vol. 10, 1661, 08.2019, p. 1-8.
  6. Blagovechtchenski, Evgeny; Gnedykh, Daria; Kurmakaeva, Diana; Mkrtychian, Nadezhda; Kostromina, Svetlana; Shtyrov, Yury. Transcranial direct current stimulation (Tdcs) of wernicke’s and broca’s areas in studies of language learning and word acquisition. Journal of Visualized Experiments, Vol. 2019, No. 149, e59159 , 07.2019, p. 1-14.
  7. Shtyrov, Yury; Kirsanov, Alexander; Shcherbakova, Olga. Explicitly Slow, Implicitly Fast, or the Other Way Around? Brain Mechanisms for Word Acquisition. Frontiers in Human Neuroscience, Vol. 13, 116, 26.04.2019.
  8. Novitskiy, Nikolay; Myachykov, Andriy; Shtyrov, Yury. Crosslinguistic interplay between semantics and phonology in late bilinguals : neurophysiological evidence. Bilingualism, Vol. 22, No. 2, 03.2019, p. 209-227.
  9. Kimppa, Lilli; Shtyrov, Yury; Hut, Suzanne C.A.; Hedlund, Laura; Leminen, Miika; Leminen, Alina Acquisition of L2 morphology by adult language learners. Cortex, 01.01.2019.
  10.  Shcherbakova, O. V.; Filippova, M. G.; Shtyrov, Y. Y. Error-related negativity as a marker of unconscious sensitivity to ambiguity. Zhurnal Vysshei Nervnoi Deyatelnosti Imeni I.P. Pavlova, Vol. 69, No. 3, 2019, p. 334-338.
  11. Scheepers, Christoph; Galkina, Anastasia; Shtyrov, Yury; Myachykov, Andriy. Hierarchical structure priming from mathematics to two- and three-site relative clause attachment. Cognition, Vol. 189, 2019, p. 155-166.
  12. Filippova, M. G.; Shcherbakova, O. V.; Shtyrov, Y. Y. It is not what you think it is: Erp correlates of verbal and non-verbal ambiguity processing. Zhurnal Vysshei Nervnoi Deyatelnosti Imeni I.P. Pavlova, Vol. 69, No. 2, 2019, p. 175-186.
  13. Kimppa, Lilli; Shtyrov, Yury; Partanen, Eino; Kujala, Teija. Impaired neural mechanism for online novel word acquisition in dyslexic children. Scientific Reports, Vol. 8, No. 1, 12779, 01.12.2018.
  14. Vukovic, Nikola; Fardo, Francesca; Shtyrov, Y. U.R.Y. When words burn – Language processing differentially modulates pain perception in typical and chronic pain populations. Language and Cognition, Vol. 10, No. 4, 12.2018, p. 626-640.
  15. Koshkin, Roman; Shtyrov, Yury; Myachykov, Andriy; Ossadtchi, Alex Testing the efforts model of simultaneous interpreting : An ERP study. PLOS ONE, Vol. 13, No. 10, 0206129, 24.10.2018, p. e0206129.
  16. Stupina, Ekaterina; Myachykov, Andriy; Shtyrov, Yury Automatic Lexical Access in Visual Modality : Eye-Tracking Evidence. Frontiers in Psychology, Vol. 9, 1847, 02.10.2018, p. 1847.
  17. Martin-Luengo, Beatriz; Shtyrov, Yury; Luna, Karlos; Myachykov, Andriy. Different answers to different audiences : effects of social context on the accuracy-informativeness trade-off. Memory, Vol. 26, No. 7, 08.2018, p. 993-1007.
  18. Pokhoday, Mikhail; Scheepers, Christoph; Shtyrov, Yury; Myachykov, Andriy. Motor (but not auditory) attention affects syntactic choice. PLOS ONE, Vol. 13, No. 4, e0195547, 01.04.2018.
  19. Gosselke Berthelsen, Sabine; Horne, Merle; Brännström, K. Jonas; Shtyrov, Yury; Roll, Mikael. Neural processing of morphosyntactic tonal cues in second-language learners. Journal of Neurolinguistics, Vol. 45, 01.02.2018, p. 60-78.
  20. Ulanov, M. A.; Stroganova, T. A.; Shtyrov, Y. Y. Language rehabilitation in chronic post-stroke aphasia: A neuroscientific perspective. Psychology, Journal of the Higher School of Economics, Vol. 15, No. 2, 01.01.2018, p. 232-245.
  21. Partanen, Eino; Leminen, Alina; Cook, Clare; Shtyrov, Yury. Formation of neocortical memory circuits for unattended written word forms: neuromagnetic evidence. Scientific Reports, Vol. 8, No. 1, 15829, 2018.
  22. Gansonre, Christelle; Højlund, Andreas; Leminen, Alina; Bailey, Christopher; Shtyrov, Yury Task-free auditory EEG paradigm for probing multiple levels of speech processing in the brain. Psychophysiology, Vol. 55, No. 11, e13216, 2018.
  23. Dagaev, Nikolay; Shtyrov, Yury; Myachykov, Andriy.The role of executive control in the activation of manual affordances. Psychological Research, Vol. 81, No. 6, 11.2017, p. 1110-1124.
  24. Partanen, Eino; Leminen, Alina; de Paoli, Stine; Bundgaard, Anette ; Kingo, Osman Skjold; Krøjgaard, Peter; Shtyrov, Yury. Flexible, rapid and automatic neocortical word form acquisition mechanism in children as revealed by neuromagnetic brain response dynamics. NeuroImage, Vol. 155, 15.07.2017, p. 450-459.
  25. Vukovic, Nikola; Feurra, Matteo; Shpektor, Anna; Myachykov, Andriy; Shtyrov, Yury. Primary motor cortex functionally contributes to language comprehension: An online rTMS study. Neuropsychologia, Vol. 96, 01.02.2017, p. 222-229.
  26. Vukovic, Nikola; Shtyrov, Yury Cortical networks for reference-frame processing are shared by language and spatial navigation systems. NeuroImage, Vol. 161, 2017, p. 120-133.
  27. Shtyrov, Yury; Lenzen, Maria.First-pass neocortical processing of spoken language takes only 30 msec : Electrophysiological evidence. Cognitive Neuroscience, Vol. 8, No. 1, 2017, p. 24-38.
  28. Chennu, Srivas; Noreika, Valdas; Gueorguiev, David; Shtyrov, Yury; Bekinschtein, Tristan A; Henson, Richard. Silent Expectations : Dynamic Causal Modeling of Cortical Prediction and Attention to Sounds That Weren’t. The Journal of neuroscience : the official journal of the Society for Neuroscience, Vol. 36, No. 32, 10.08.2016, p. 8305-16.
  29. Kimppa, Lilli; Kujala, Teija; Shtyrov, Yury Individual language experience modulates rapid formation of cortical memory circuits for novel words. Scientific Reports, Vol. 6, 30227, 22.07.2016.
  30. Leminen, Alina; Kimppa, Lilli; Leminen, Miika M; Lehtonen, Minna; Mäkelä, Jyrki P; Shtyrov, Yury. Acquisition and consolidation of novel morphology in human neocortex : A neuromagnetic study. Cortex, Vol. 83, 09.07.2016, p. 1-16.
  31. Mohr, Bettina; MacGregor, Lucy J; Difrancesco, Stephanie; Harrington, Karen; Pulvermüller, Friedemann; Shtyrov, Yury. Hemispheric contributions to language reorganisation : An MEG study of neuroplasticity in chronic post stroke aphasia. Neuropsychologia, 07.04.2016.
  32. Hanna, Jeff; Shtyrov, Yury; Williams, John; Pulvermüller, Friedemann Early neurophysiological indices of second language morphosyntax learning. Neuropsychologia, Vol. 82, 02.2016, p. 18-30.
  33. Egorova, Natalia; Shtyrov, Yury; Pulvermüller, Friedemann. Brain basis of communicative actions in language. NeuroImage, Vol. 125, 15.01.2016, p. 857-67.
  34. Luna, Karlos; Martín-Luengo, Beatriz; Shtyrov, Yury; Myachykov, Andriy.Judgments of Learning for Words in Vertical Space. Frontiers in Psychology, Vol. 7, 2016, p. 1894. 
  35. Shtyrov, Yury; MacGregor, Lucy J. Near-instant automatic access to visually presented words in the human neocortex : neuromagnetic evidence. Scientific Reports, Vol. 6, 2016, p. 26558.
  36. Moseley, Rachel L; Correia, Marta M; Baron-Cohen, Simon; Shtyrov, Yury; Pulvermüller, Friedemann; Mohr, Bettina. Reduced Volume of the Arcuate Fasciculus in Adults with High-Functioning Autism Spectrum Conditions. Frontiers in Human Neuroscience, Vol. 10, 2016, p. 214.
  37. Kimppa, Lilli; Kujala, Teija; Leminen, Alina; Vainio, Martti; Shtyrov, Yury. Rapid and automatic speech-specific learning mechanism in human neocortex. NeuroImage, Vol. 118, 09.2015, p. 282-91.
  38. Roll, Mikael; Söderström, Pelle; Mannfolk, Peter; Shtyrov, Yury; Johansson, Mikael; van Westen, Danielle; Horne, Merle. Word tones cueing morphosyntactic structure : Neuroanatomical substrates and activation time-course assessed by EEG and fMRI. Brain and Language, Vol. 150, 17.08.2015, p. 14-21.
  39. Ylinen, Sari; Nora, Anni; Leminen, Alina; Hakala, Tero; Huotilainen, Minna; Shtyrov, Yury; Mäkelä, Jyrki P; Service, Elisabet. Two distinct auditory-motor circuits for monitoring speech production as revealed by content-specific suppression of auditory cortex. Cerebral Cortex, Vol. 25, No. 6, 06.2015, p. 1576-86.
  40. MacGregor, Lucy J; Difrancesco, Stephanie; Pulvermüller, Friedemann; Shtyrov, Yury; Mohr, Bettina. Ultra-rapid access to words in chronic aphasia: the effects of intensive language action therapy (ILAT). Brain Topography, Vol. 28, No. 2, 03.2015, p. 279-91.
  41. Whiting, Caroline; Shtyrov, Yury; Marslen-Wilson, William. Real-time functional architecture of visual word recognition. Journal of Cognitive Neuroscience (Online), Vol. 27, No. 2, 02.2015, p. 246-65.
  42. Moseley, R. L.; Shtyrov, Yury; Mohr, B.; Lombardo, M. V.; Baron-Cohen, S.; Pulvermuller, F. Lost for emotion words: what motor and limbic brain activity reveals about autism and semantic theory. NeuroImage, Vol. 104, No. 1, 2015, p. 413-422.
  43. Shtyrov, Yury Y; Stroganova, Tatiana A. When ultrarapid is ultrarapid: on importance of temporal precision in neuroscience of language. Frontiers in Human Neuroscience, Vol. 9, No. 576, 2015.
  44. Vukovic, Nikola; Shtyrov, Yury Cortical motor systems are involved in second-language comprehension : Evidence from rapid mu-rhythm desynchronisation. NeuroImage, Vol. 102, No. P2, 05.11.2014, p. 695-703.
  45. Moseley, Rachel L.; Pulvermüller, Friedemann; Mohr, Bettina; Lombardo, Michael V.; Baron-Cohen, Simon; Shtyrov, Yury. Brain routes for reading in adults with and without autism : EMEG evidence. Journal of Autism and Developmental Disorders, Vol. 44, No. 1, 01.01.2014, p. 137-153.
  46. Hanna, Jeff; Mejias, Sandrine; Schelstraete, Marie Anne; Pulvermüller, Friedemann; Shtyrov, Yury; van der Lely, Heather K J. Early activation of Broca’s area in grammar processing as revealed by the syntactic mismatch negativity and distributed source analysis. Cognitive Neuroscience, Vol. 5, No. 2, 01.01.2014, p. 66-76.
  47. Egorova, Natalia; Pulvermul̈ler, Friedemann; Shtyrov, Yury. Neural dynamics of speech act comprehension : An MEG study of naming and requesting. Brain Topography, Vol. 27, No. 3, 01.01.2014, p. 375-392.
  48. Shtyrov, Yury; Butorina, Anna; Nikolaeva, Anastasia; Stroganova, Tatiana Automatic ultrarapid activation and inhibition of cortical motor systems in spoken word comprehension. Proceedings of the National Academy of Sciences of the United States of America, Vol. 111, No. 18, 2014, p. E1918-23.
  49. Leminen, Alina; Leminen, Miika; Kujala, Teija; Shtyrov, Yury. Neural dynamics of inflectional and derivational morphology processing in the human brain. Cortex, Vol. 49, No. 10, 01.10.2013, p. 2758-71.
  50. Macgregor, Lucy J; Shtyrov, Yury. Multiple routes for compound word processing in the brain : Evidence from EEG. Brain and Language, Vol. 126, No. 2, 22.06.2013, p. 217-229.
  51. Bakker, Iske; Macgregor, Lucy J; Pulvermüller, Friedemann; Shtyrov, Yury. Past tense in the brain’s time : neurophysiological evidence for dual-route processing of past-tense verbs. NeuroImage, Vol. 71, 01.05.2013, p. 187-95.
  52. Shtyrov, Yury; Goryainova, Galina; Tugin, Sergei; Ossadtchi, Alexey; Shestakova, Anna Automatic processing of unattended lexical information in visual oddball presentation : neurophysiological evidence. Frontiers in Human Neuroscience, Vol. 7, 2013, p. 421.
  53. Egorova, Natalia; Shtyrov, Yury; Pulvermüller, Friedemann. Early and parallel processing of pragmatic and semantic information in speech acts : neurophysiological evidence. Frontiers in Human Neuroscience, Vol. 7, 2013, p. 86.
  54. Myachykov, Andriy; Scheepers, Christoph; Shtyrov, Yury. Interfaces between language and cognition. Frontiers in Psychology, Vol. 4, 2013, p. 258.
  55. Whiting, Caroline M; Marslen-Wilson, William D; Shtyrov, Yury. Neural dynamics of inflectional and derivational processing in spoken word comprehension : laterality and automaticity. Frontiers in Human Neuroscience, Vol. 7, 2013, p. 759.
  56. Moseley, Rachel L; Pulvermüller, Friedemann; Shtyrov, Yury. Sensorimotor semantics on the spot : brain activity dissociates between conceptual categories within 150 ms. Scientific Reports, Vol. 3, 2013, p. 1928.
  57. Shtyrov, Yury; Smith, M. L.; Horner, A. J.; Henson, R.; Nathan, P. J.; Bullmore, E. T.; Pulvermuller, F. Attention to language: novel MEG paradigm for registering involuntary language processing in the brain. Neuropsychologia, Vol. 50, 2012, p. 2605-2616.
  58. Nicol, R. M.; Chapman, S. C.; Vertes, P. E.; Nathan, P. J.; Smith, M. L.; Shtyrov, Yury; Bullmore, E. T. Fast reconfiguration of high-frequency brain networks in response to surprising changes in auditory input. Journal of Neurophysiology, Vol. 107, 2012, p. 1421-1430.
  59. Shtyrov, Yury Neural bases of rapid word learning. Neuroscientist, Vol. 18, 2012, p. 312-319.
  60. Holland, R.; Brindley, L.; Shtyrov, Yury; Pulvermuller, F.; Patterson, K. They played with the trade: MEG investigation of the processing of past tense verbs and their phonological twins. Neuropsychologia, Vol. 50, 2012, p. 3713-3720.
  61. MacGregor, L. J.; Pulvermuller, F.; van Casteren, M.; Shtyrov, Yury. Ultra-rapid access to words in the brain. Nature Communications, Vol. 3, 2012, p. 711.
  62. Boulenger, V.; Shtyrov, Yury; Pulvermuller, F. When do you grasp the idea? MEG evidence for instantaneous idiom understanding. NeuroImage, Vol. 59, 2012, p. 3502-3513.
  63. Shtyrov, Yury; Kimppa, Lilli; Pulvermüller, Friedemann; Kujala, Teija.Event-related potentials reflecting the frequency of unattended spoken words : a neuronal index of connection strength in lexical memory circuits? NeuroImage, Vol. 55, No. 2, 15.03.2011, p. 658-68.
  64. Shtyrov, Yury; Kimppa, L.; Pulvermuller, F.; Kujala, T. Event-related potentials reflecting the frequency of unattended spoken words: a neuronal index of connection strength in lexical memory circuits? NeuroImage, Vol. 55, 2011, p. 658-668.
  65. Shtyrov, Yury. Fast mapping of novel word forms traced neurophysiologically. Frontiers in Psychology, Vol. 2, 2011, p. 340.
  66. Alexandrov, A. A.; Boricheva, D. O.; Pulvermuller, F.; Shtyrov, Yury. Strength of word-specific neural memory traces assessed electrophysiologically. PLoS One. Vol. 6, 2011, p. e22999.
  67. Shtyrov, Yury Automaticity and attentional control in spoken language processing : Neurophysiological evidence. Mental Lexicon, Vol. 5, No. 2, 23.12.2010, p. 255-276.
  68. Cappelle, B.; Shtyrov, Yury; Pulvermuller, F. Heating up or cooling up the brain? MEG evidence that phrasal verbs are lexical units. Brain and Language, Vol. 115, 2010, p. 189-201.
  69. Shtyrov, Yury; Kujala, T.; Pulvermuller, F. Interactions between language and attention systems: early automatic lexical processing? Journal of Cognitive Neuroscience (Online), Vol. 22, 2010, p. 1465-1478.
  70. Shtyrov, Yury; Nikulin, V. V.; Pulvermuller, F. Rapid cortical plasticity underlying novel word learning. The Journal of neuroscience : the official journal of the Society for Neuroscience, Vol. 30, 2010, p. 16864-16867.
  71. Vestergaard, M. D.; Haden, G. P.; Shtyrov, Yury; Patterson, R. D.; Pulvermuller, F.; Denham, S. L.; Sziller, I.; Winkler, I. Auditory size-deviant detection in adults and newborn infants. Biological Psychology, Vol. 82, 2009, p. 169-175.
  72. Carlyon, R. P.; Deeks, J. M.; Shtyrov, Yury; Grahn, J.; Gockel, H. E.; Hauk, O.; Pulvermuller, F. Changes in the perceived duration of a narrowband sound induced by a preceding stimulus Journal of experimental psychology. Human perception and performance, Vol. 35, 2009, p. 1898-1912.
  73. Garagnani, M.; Shtyrov, Yury; Pulvermuller, F. Effects of attention on what is known and what is not: MEG evidence for functionally discrete memory circuits. Frontiers in Human Neuroscience, Vol. 3, 2009, p. 10.
  74. Pulvermuller, F.; Shtyrov, Yury. Spatiotemporal signatures of large-scale synfire chains for speech processing as revealed by MEG. Cerebral Cortex, Vol. 19, 2009, p. 79-88.
  75. Pulvermuller, F.; Shtyrov, Yury; Hauk, O. Understanding in an instant: neurophysiological evidence for mechanistic language circuits in the brain. Brain and Language, Vol. 110, 2009, p. 81-94.
  76. Shtyrov, Yury; Osswald, K.; Pulvermuller, F. Memory traces for spoken words in the brain as revealed by the hemodynamic correlate of the mismatch negativity. Cerebral Cortex, Vol. 18, 2008, p. 29-37.
  77. Pulvermuller, F.; Shtyrov, Yury; Hasting, A. S.; Carlyon, R. P. Syntax as a reflex: neurophysiological evidence for early automaticity of grammatical processing. Brain and Language, Vol. 104, 2008, p. 244-253.
  78. Shtyrov, Yury; Pulvermüller, Friedmann Language in the mismatch negativity design : Motivations, benefits, and prospects. Journal of psychophysiology, Vol. 21, No. 3-4, 01.12.2007, p. 176-187.
  79. Shtyrov, Yury; Pulvermuller, F. Early MEG activation dynamics in the left temporal and inferior frontal cortex reflect semantic context integration. Journal of Cognitive Neuroscience, Vol. 19, 2007, p. 1633-1642.
  80. Pulvermuller, F.; Shtyrov, Yury. Language outside the focus of attention: the mismatch negativity as a tool for studying higher cognitive processes. Progress in Neurobiology, Vol. 79, 2006, p. 49-71.
  81. Pulvermuller, F.; Huss, M.; Kherif, F.; Moscoso del Prado Martin, F.; Hauk, O.; Shtyrov, Yury. Motor cortex maps articulatory features of speech sounds. Proceedings of the National Academy of Sciences of the United States of America, Vol. 103, 2006, p. 7865-7870.
  82. Hauk, O.; Shtyrov, Yury; Pulvermuller, F. The sound of actions as reflected by mismatch negativity: rapid activation of cortical sensory-motor networks by sounds associated with finger and tongue movements. The European journal of neuroscience, Vol. 23, 2006, p. 811-821.
  83. Pulvermuller, F.; Shtyrov, Yury; Ilmoniemi, R. J.; Marslen-Wilson, W. D. Tracking speech comprehension in space and time. NeuroImage, Vol. 31, 2006, p. 1297-1305.
  84. Uther, M.; Kujala, A.; Huotilainen, M.; Shtyrov, Yury; Naatanen, R. Training in Morse code enhances involuntary attentional switching to acoustic frequency: Evidence from ERPs. Brain Research, Vol. 1073-1074, 2006, p. 417-424.
  85. Pulvermuller, F.; Shtyrov, Yury; Ilmoniemi, R. Brain signatures of meaning access in action word recognition. Journal of Cognitive Neuroscience, Vol. 17, 2005, p. 884-892.
  86. Shtyrov, Yury; Pihko, E.; Pulvermuller, F. Determinants of dominance: is language laterality explained by physical or linguistic features of speech? NeuroImage, Vol. 27, 2005, p. 37-47.
  87. Kujala, T.; Shtyrov, Yury; Winkler, I.; Saher, M.; Tervaniemi, M.; Sallinen, M.; Teder-Salejarvi, W.; Alho, K.; Reinikainen, K.; Naatanen, R. Long-term exposure to noise impairs cortical sound processing and attention control. Psychophysiology, Vol. 41, 2004, p. 875-881.
  88. Pulvermuller, F.; Shtyrov, Yury; Kujala, T.; Naatanen R. Word-specific cortical activity as revealed by the mismatch negativity. Psychophysiology. Vol. 41, 2004, p. 106-112.
  89. Pulvermuller, F.; Shtyrov, Yury. Automatic processing of grammar in the human brain as revealed by the mismatch negativity. NeuroImage, Vol. 20, 2003, p. 159-172.
  90. Shtyrov, Yury; Pulvermuller, F.; Naatanen, R.; Ilmoniemi, R. J. Grammar processing outside the focus of attention: an MEG study. Journal of Cognitive Neuroscience, Vol. 15, 2003, p. 1195-1206.
  91. Kujala, A.; Huotilainen, M.; Uther, M.; Shtyrov, Yury; Monto, S.; Ilmoniemi, R. J.; Naatanen, R. Plastic cortical changes induced by learning to communicate with non-speech sounds. NeuroReport, Vol. 14, 2003, p. 1683-1687.
  92. Pulvermuller, F.; Shtyrov, Yury; Ilmoniemi, R. Spatiotemporal dynamics of neural language processing: an MEG study using minimum-norm current estimates. NeuroImage, Vol. 20, 2003, p. 1020-1025.
  93. Micheyl, C.; Carlyon, R. P.; Shtyrov, Yury; Hauk, O.; Dodson, T.; Pullvermuller, F. The neurophysiological basis of the auditory continuity illusion: a mismatch negativity study Journal of Cognitive Neuroscience, Vol. 15, 2003, p. 747-758.
  94. Palva, S.; Palva, J. M.; Shtyrov, Yury; Kujala, T.; Ilmoniemi, R. J.; Kaila, K.; Naatanen, R.Distinct gamma-band evoked responses to speech and non-speech sounds in humans. The Journal of neuroscience : the official journal of the Society for Neuroscience, Vol. 22, 2002, p. RC211.
  95. Shtyrov, Yury; Pulvermuller, F. Memory traces for inflectional affixes as shown by mismatch negativity. The European journal of neuroscience, Vol. 15, 2002, p. 1085-1091.
  96. Shtyrov, Yury; Pulvermuller, F. Neurophysiological evidence of memory traces for words in the human brain. NeuroReport, Vol. 13, 2002, p. 521-525.
  97. Pulvermuller, F.; Kujala, T.; Shtyrov, Yury; Simola, J.; Tiitinen, H.; Alku, P.; Alho, K.; Martinkauppi, S.; Ilmoniemi, R. J.; Naatanen, R. Memory traces for words as revealed by the mismatch negativity. NeuroImage, Vol. 14, 2001, p. 607-616.
  98. Shtyrov, Yury; Kujala, T.; Lyytinen, H.; Ilmoniemi, R. J.; Naatanen, R. Auditory cortex evoked magnetic fields and lateralization of speech processing. NeuroReport, Vol. 11, 2000, p. 2893-2896.
  99. Shtyrov, Yury; Kujala, T.; Palva, S.; Ilmoniemi, R. J.; Naatanen, R. Discrimination of speech and of complex nonspeech sounds of different temporal structure in the left and right cerebral hemispheres. NeuroImage, Vol. 12, 2000, p. 657-663.
  100. Shtyrov, Yury; Kujala, T.; Lyytinen, H.; Kujala, J.; Ilmoniemi, R. J.; Naatanen, R. Lateralization of speech processing in the brain as indicated by mismatch negativity and dichotic listening. Brain and Cognition, Vol. 43, 2000, p. 392-398.
  101. Pulvermüller, Friedemann; Teija, Kujala; Shtyrov, Yury; Simola, Jaana; Alku, Paavo; Alho, Kimmo; Näätänen, Risto. Phonological and lexico-semantic factors influence the mismatch negativity (MMN). NeuroImage, Vol. 9, No. 6 PART II, 01.12.1999.
  102. Shtyrov, Yury; Kujala, T.; Ilmoniemi, R. J.; Naatanen, R. Noise affects speech-signal processing differently in the cerebral hemispheres. NeuroReport, Vol. 10, 1999, p. 2189-2192.
  103. Shtyrov, Yury; Kujala, T.; Ahveninen, J.; Tervaniemi, M.; Alku, P.; Ilmoniemi, R. J.; Naatanen, R. Background acoustic noise and the hemispheric lateralization of speech processing in the human bra magnetic mismatch negativity study. Neuroscience Letters, Vol. 251, 1998, p. 141-144.
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